Notes on Dictyolathys Maculata Banks (Araneæ: Dictynidæ)

Psyche: A Journal of Entomology, Mar 2019

Elizabeth B. Bryant

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Notes on Dictyolathys Maculata Banks (Araneæ: Dictynidæ)

International Journal of Notes on Dictyolathys maculata BY ELIZABETH B. BRYANT 0 0 Museum of Comparative ZoiSlogy , Cambridge, Mass - Recently in examining some material from Raleigh, North Carolina, collected in January 1941, two females were found described .as Dictyo!athys maculata Banks. This genus, erected by Banks in 1900 , was based on females from Mobile, Alabama, and Meridian, Mississippi, and was separ.ated from Lathys by the strongly procurved anterior eye row. In 1903, Simon, who had never seen a specimen, pl.aced the genus as .a synonym of Lathys and was followed by Emerton, in 1913 . He described and figured the male from specimens collected May 1, 1912 .at Lakehurst, New Jersey. At the end of the description, Emert.on says, "In the original description of Dictyolathys maculata, Banks says that the front middle eyes can be indistinctly seen but I have not been able to find. them in the Lakehurst specimens nor in the (Banks) type specimen from Alabama. They appear to have only six eyes in two groups as in Scotolathys pallida." Today with the sharp light from a condenser, the small anterior median eyes seen by Banks can be distinctly placed both in the type and the specimens used by Emerton, as well as in the recent material from Raleigh, so that the Banks genus Dictyolathys can no longer be considered a synonym either of Lathys or Scotolathys. For the benefit of those who do not have ready access to the original description of Dictyolathys Banks, I quote verbatim as foll.ows: "Much like Dictyna, but .apparently six-eyed, three in a grotap each side but the A.M.E..are present, although very small, and situated close to and a little higher than the A.S.E. Head not much elevated; legs of moderate length, no spines, but very hairy; .accessory spinning organs like Dictyna." A more detailed description of the genus can now be written to include other char.acters not used by Banks in the original description. Genus Dictyolathys Banks Cephalothorax moderately high, no thoracic groove; eight eyes, lateral eyes on a low tubercle, a.m.e, very small on the posterior slope of the tubercle, widely separated, so that the anterior row of eyes is strongly procurved, posterior row slightly procurved, p.l.e, largest of the eight; clypeus equals a radius of a.l.e.; labium triangular, as long as wide; sternum convex, almost as wide as long, ending in a lobe between fourth coxa; legs varying little in length, fourth pair longest, no spines, dorsal row of trichobothria on fourth tibia; calamistrum confined to basal half of fourth metatarsus. Dictyolathys is separated from Lathys by the strongly procurved anterior eye row, .and the p.l.e, larger than the p.m.e., and from Scotolathys by the presence of small a.m.e..and the large p.l.e. The genus Lathys Simon, 1884, was based on the species humilis Blackwall, common to northern Europe. This has eight eyes, with the anterior row straight, eyes contiguous, a.m.e. very small, posterior row slightly procurved, eyes equal and usually equidistant. The genus Scotolathys Simon, 1884, was erected for a species,. (simplex) from Algiers, with but six eyes, the .a.l.e. the largest of the eight. In both Les Histoire Naturelle des Araignes, 1903, 2, p. 977 and Les Arachnides de France, 1914, 6, p. 62, Simon has placed Lathys heterophthalma Kulczynski, 1891, in the genus Scotolathys but states that the a.m.e..almost obliterated. No specimens of this species are in the museum collection; but in the original description, " Oculorum; series posterior paullo procurva, oculi magni, inter se subaquales et spatiis parum inaqualibus, circiter radium equantibus remoti; series anterior subrecta, oculi valde inaquales, mediorum diameter radio lateralium minor, oculi laterales posticis subquales, medii inter se late (plus quam diametro sua) distantes, lateralibus v.alde approximate." Having regard to this and the figures, it would seem that Kulczynski was correct in placing the species in the genus Lathys. Dictyolathys maculata Banks Cephalothorax moderately convex, cephalic portion level, with a median row .of very long bristles from near the posterior margin to eyes, thor.acic portion falls rapidly to posterior margin; eyes in two groups that do not cover the anterior margin, anterior row strongly procurved, lateral eyes on .a tubercle, a.l.e. separated by a diameter, a.m.e, very small, .on posterior slope of tubercle, posterior row procurved, p.l.e, largest of the eight, ::I:?;::;?.:?.":".":??.. \UI "! \ ": :?"::":":i.:".?:.:,.. "--___ .:..f.::..::...:;.i:i:r FtG. 1. Dictyolathys maculata Banks. A, eyes; B, epigynum. separated from a.l.e, by .a line and from p.m.e, by less than a radius, p.m.e, separated by about a diameter; quadrangle of median eyes much narrower in front and as high as wide behind; clypeus .as high as radius of .a.l.e.; mandibles small, verticle; labium triangular, with tip black, not fused to sternum; maxillae inclined over labium with tips almost touching; sternum pale, strongly convex, only slightly longer than wide, ending in .a lobe between IV coxe, IV coxe widely separated; abdomen, pale, with a small median basal black spot, followed by five pairs of diagonal black spots, sides with black streaks, venter pale; legs, pale, relatively short, varying little in length, but the IV pair longest, no spines, a dorsal row of trichobothria on IV tibia; calamistrum of curved bristles, less, than the diameter of the joint in length, on basal half of IV metatarsus; epigynum large for the size of the spider, chitinized area wider than long, with openings widely separated, and a pale depressed area just anterior to openings. Dictyolathys maculata Banks has been reported by Bishop and Crosby in 1926 , from Madison and Raleigh, North Carolina and by Gertsch and Mulaik in 1940, from Houston, Harris Co., Texas. Material in the Museum of Compartive ZoiSlogy Collection: Holotype Ala.; Mobile, Banks Coll. 2 N. J.; Lakehurst, 1 May 1912, Emert.on Coll. 2 N. C.; Raleigh, oak woods, 23 January 1941, (Wray). In 1904, Banks described Dictyolathys caliornica from P,alo Alto, California, also from a female. Unfortunately, this specimen cannot be found in the museum collection, but from the description and the figures, it probably does not belong to the genus Dictyolathys since the small a.m.e, are only a diameter apart; the anterior row is almost straight as in Lathys and the other eyes are subequal. Peptides Advance s in ht p:/ w Virol og y Hindawi Publishing Corporation Hindawi Publishing Corporation ht p:/ Submit your manuscr ipts BioMed Hindawi Publishing Corporation ht p:/ Research International Stem Cells International Hindawi Publishing Corporation ht p:/ Zoology International Journal of Journal of Signal Hindawi Publishing Corporation ht p:/ Transduction Genetics Research International Hindawi Publishing Corporation ht p:/ Anatomy Research International Research International Advances in Bioinformatics Hindawi Publishing Corporation ht p:/ Hindawi Publishing Corporation ht p:/ Enzyme Research International Journal of Journal of International Journal of Genomics Hindawi Publishing Corporation ht p:/ Journal of Nucleic Acids The Scientiifc World Journal Banks , Nathan. 1900 . Some Arachnida from Alabama . Proc. Philadelphia Acad ., pp. 529 - 543 . 1904. Some Arachnida from California . Proc. California Acad., 3rd ser., 3 , pp. 331 - 369 , pls. 38 - 41 . Chyzer et Kulczynski. 1892 . Aranee Hungarian, vol. 1 , p. 161 . Emerton , J. H. 1913 . New and Rare Spiders from within fifty miles of New York City . Bull. American Mus. Nat. Hist., 32 , pp. 255 - 260 , pl. 48 . Simon , Eugene. 1913 . Histoire Naturelle des Araignes, 2 , p. 977 . 1914 . Arachnides de France, 6 , p. 62 . Bishop , S. C. and Crosby , C. R. 1926 . Notes on the Spiders of the Southeastern United States with Descriptions of New Species . Jour. Elisha Mitchell Sci. Soc., 41 , pp. 165 - 212 , pls. 20 - 25 .

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Elizabeth B. Bryant. Notes on Dictyolathys Maculata Banks (Araneæ: Dictynidæ), Psyche: A Journal of Entomology, DOI: 10.1155/1943/23428