New and Little Known Species of Tegenaria (Araneida Agelenidae)

Psyche: A Journal of Entomology, Mar 2019

Harriet Exline

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New and Little Known Species of Tegenaria (Araneida Agelenidae)

International Journal of Little Known Species of Tegenaria Psyche 0 0 University of Washington , USA (Araneida Agelenidae) BY HARRIET' EXLINE Three new species o.f Tegenaria have been found in the Pacific Northwest and are described in this paper. Two other species, Tegenaria nana Simon and Tegvnaria californica Banks, are included because they are quite common in the Northwest, and because their original descriptions included no figures. The most, common Tegenaria represented in this locality is the cosmopolitan species Tegenaria derhami (Scop.), a description of which is omitted because it is adequately described and figured elsewhere (Emerton: Common Spiders, 1902, p. 96; ibid: New Eng. Spiders, Trans. Conn. Acad., Vol. VIII, 1889, p. 29, f. 6.). Notes of appreciation are expressed to Miss Elizabeth B. Bryant of the Museum of Compa.rative ZoSlogy f.or identilying specimens of Tegenaria, and to Drs. Melville H. Hatch and Robert C. Miller of 'the University of Washington for their help and inspiration, and especially to Professor Trevor Kincaid for the loan of his collection of spiders. - NEW AND LITTLE KNOWN SPECIES OF TEGENARIA Tegenaria gigantea Chamberlin and Ivie, 1935, Bull. Univ. Utah, vol. 26, p. 31, pl. XIII, f. 106. MALE" Length 15 mm., with extremely long, slender and hairy legs. Cephalothorax reddish yellow, becoming quite reddish on cephalic part. and almost black around and in front of the eyes. The cephalic part is roun.ded, highest in the center, the sides are almost parallel, and the entire cephalic region is raised and sharply separated from the thoracic part which is rather flat and almost circular in outline. The chelicerae and mouthparts are almost black, somewhat geniculate, with prominent yellow condyles. The eyes are about equal in size, although the posterior eyes are not quite so large as the anterior eyes, the posterior eyes in a slightly procurved row, well and equally separated; anterior eyes in an almost straight row (rom a dorsal view) and closer together. Anterior median eyes the radius of one of them apart, and not quite so. 2ar 2rom the anterior lateral eyes. Legs tannish yellow except femora of first pair which are brown; the first leg measures 4.5 cm. The abdomen is slender, and yellow with many ,dark gray markings; the area in the a.nterior median region is unmarked. The. hind spinnerets are long, with the apical segments a little longer than he basal segment. The palpus is slender with the cymbium well clothed with long hairs. The tibia of the palpus is slightly sw.ollen distally and has two large toothshaped apophyses extending latero-anteriorly on the lateral side.. The embolus of the bulb is. slender and arises from the inner distal margin, curving la.terally to end in a projecting conductor in the lateral posterior region. Male, collected at Nanaimo, B. C. by Professor Kincaid, Sept. 8, 1934. (Exline Coll.) This species nearest Tegenaria derhami Scop. rom which it is separated by its much grea.ter length and by the structure of the bulb o the palpus. ?Teena,ria quadrata n. sp. (Fi$. 4) FEMALE: Length 5.6 mm. Cephalothorax brownish yellow with a few irregular gray marks including a narrow marginal line,, a.nd a. row of spots, between this and the median furrow opposite the legs, and a large median gray area. Chelicerae and legs same color as the cephalo.horax, legs banded with gray. Eyes la.rge and quite widely separated; posterior eyes in a straight, row equally sPaced and equal in size; anterior median eyes smaller than other eyes separated by a distance once and a half as great as their diameter, nearer the a.nterior lateral, eyes,; thus the rectangle formed by the median eyes is large and perfect, longer than wide.. Abdomen globose with a gray-white background a.nd many darker gray markings. The apical segment of the hind spinnerets is a little shorter than the basal segment. The area ,of he epigynum is raised and heavily Little Kown Species of Tegenaria chitinized; on the anterior margin is a small depression which obviously pores leading into. the spermathecal ducts; these ducts are indistinctly seen through the chitin extending from Che depression a.lmost to the genital urrow in the midline. The spermathecae can be indistinctly seen in the anterior lateral parts of the epigynum. HOLOTYPE: Female., collected a Sea.ttle, Wash., by Proess.or Kincaid. (Exline Coll.) Closest to Tegenaria californica Banks, but separa.ted by the much grea.ter distance between the anterior median eyes, and the large chitinized area of the epigynum which is negligible in T. californica.. Tegenaria magnacava n. sp. (Fig. 5) FEMALE: Length 11 mm. Cephalothorax light reddish yellow with grayish streaks on the sides, cephalic part only a little darker than the thoracic; cephalothorax moderately rounded with no grea.t demarcation between cephalic part and thoracic. Chelicerae quite geniculate in front, red in color. Legs well clothed with hair, moderately long and tannish or reddish yellow. Eyes. of equal size, those of posterior row so.mewhat procurved, widely .a.nd equally spaced. Eyes of anterior row almost equally spaced, but slightly nearer each other tha.n those of posterior row. Abdomen with a light yellowish gray background and mottled a.nd checked with dark gray markings. Epigynum large surrounded by a thick posterior fold of chitin and an anterior semicircular fold; the cavity formed by these, is large and deep. See fig. HOLOTYPE: Female, collected at Seattle, Wash., Nov. 20, 1930. (Exline Coll.) Similar in s.ize and physical structure to Tegenaria derhami Scop. The epigynum is entirely different in orm from that of any other species, consisting of a large cavity surrounded by chitinous folds. Tegenaria nana Simon (Fig. 1) Te.genaria nana Simon 1897, Annal. Soc. Entom. Belgique, Vol. XLI, p. 17. FEMALE: Length, 4.0-5.5 mm. Cephalothorax tannish yellow with three indefinite gra.y spots on each side of the thorax, a marginal gray band and a few gray lines on the heard part; cephalothorax rather roundly arched. Chelicerae not at all geniculate. Eyes large in proportion to size of spider; posterior eyes in nearly s,traight row, almost equally dista.nt (less than a diameter of one .of the eyes) and equal in size; eyes of anterior row very close together and median eyes considerably smaller than the later:al eyes. Legs yellow in some specimens, ora.nge or tan ringed with gray in other s,pecimens. Abdomen glob.ose, pale grayish yellow with many dark gray markings especially in the posterior half; these markings form a row of double concave arches down the cener of the back posteriorly. Apical segment, of hind spinnerets is slightly shorter than the. basal segment. The epigynum is raised, rounded off imperceptibly into the venter; in its center is a heart-shaped da,rk reddish brown chitinized area, which shows two slight impressions; these are the openings to the ducts leading to the spermathecae; posterior to this area is a slender median dark band running to. the epigastric furrow?; this dark band is caused by the ducts which lead into the sper.mathecae, which in a ew specimens are slightly visible as round balls underneath the chitin. MALE: Length, 4.0-4.5 mm. Cephalothorax very evenly rounded with cephalic grooves very shallow, yellow in color with very slight gray markings. Chelicerae, an.d relationships between the eyes. as in female; legs about the same except that they are more solidly one color, yellow, or orange brown. Abdomen slender, white background with dark gray markings numerous in the posterior part, very m?uch as in the female. Hind spinnerets with a.pical segment a little shorter than basal segment. Tibia of male palpus with two basal teeth, black in color on the lateral margins, then a deep horizontal groove which ends, in the. middle of the segment with a rather large black, flat projection directed anteriorly; on the distal .margin of the tibia it bears another lateral projection pointing anteriorly. The cymbium of the palpus is deeply rounded in its body part but anteriorly slenderizes sud.denly and has a narrow, short tip. HYPOTYPES: Male specimens, collected March 24, 1932, and Oct.. 1, 1932, Olympia, Washington (Exline Coll.). Female. specimens from Seattle, Olympia, and Beaver Creek, Oregon. Tegenaria ealiforniea Banks (Fig. 2) Tegenaria californica Banks 1896 Journ. EAt. News, Vol. 4, p. 90. FEMALE: Length 7-9 ram. Cephalothorax orange marked with black lines, and spots; legs long, hairy, ringed with dark gray. Posterior eyes in straight row, almost equidistant, laterals slightly larger than the medians; anterior eyes with medians smaller than laterals and further apart than from laterals. Abdomen large, globose, gray with light spots; posterior spinnerets with hind segment slightly shorter than basal segment. The area. of the epigyhum is decidedly convex, with a small shallow atrium in the center, in which two very small spots mark the opening of the spermathecal duct; the spermathecae show slightly through the chitin in the anterior latera:l margins; a dark ri.dge of chitin extends posteriorly from the. center of the posterior margin of the atrium to the epigastric urrow. MALE" Length 5-6.5 mm. Eyes all equal in size, eyes of posterior row and an?terior row all equidistant. Tibia of palpus with a basal, almost horizontal, apophysis and a distal pointed apophysis on the lateral margin; embolus almost encircling bulb. Distri,bution in Washington: Western part of state. Specimens from Stillaquamish River (Snohomish Co.); Lebar, Olympia Mrs.; Fort Ca.nby (Pacific Co.); Everett; Seattle; and Olympia (in Exline Coll.). 1Identifications of this and above species kindly made ,by Miss Elizabeth B. Bryant of the Museum of Comparative ZoSlogy, Harvard University. EXPLANATION OF PLATE 1. 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Harriet Exline. New and Little Known Species of Tegenaria (Araneida Agelenidae), Psyche: A Journal of Entomology, DOI: 10.1155/1936/14909